Nausicaä takes to the skies in her signature glider, engaging in aerial combat against waves of hostile forces and formidable bosses that emerge from level two onward. As she navigates the dangerous airspace, the player must master dogfighting mechanics and evasive maneuvers while using her glider's weaponry to survive increasingly difficult encounters. The gameplay centers on quick reflexes and tactical positioning as threats multiply across each stage, challenging the player to balance offense and defense in high-speed aerial duels.