Five centuries into the future, a sprawling metropolis known as the City in the Sea faces an ecological crisis when the ruthless Black Marketeers launch a systematic campaign to capture dinosaurs for mysterious purposes. Their relentless poaching has driven the creatures to desperation, transforming them from curiosities into threats that terrorize nearby settlements and jungle communities. To halt the carnage and dismantle the trafficking operation, four unlikely allies—the resourceful mechanic and spiritual leader Jack Tenrec, the intrepid explorer Hannah Dundee, the ingenious engineer Mustapha Cairo, and the enigmatic operative Mess O'Bradovich—band together to infiltrate the City in the Sea itself, where they believe the sinister hunting empire originates.