Three years after the Great Shinra War devastated the land of Soma and killed most of its plant life, humanity is working to survive and begin reforestation. An alchemist named Hiodoshi ran a project called Espgaluda, using young people as test subjects for experimental armored wings — the subjects were called Galuda. Two of them, the siblings Ageha and Tateha, escaped and lived in hiding with their mother. Espgaluda II begins when they are found.