A malevolent tyrant plagues the serene realm of Miitopia by harvesting the faces of its citizens and transplanting them onto beasts, transforming them into corrupted servants of darkness. When this calamity befalls the quiet town of Greenhorne, a mysterious divine force selects a fortuitous traveler as the chosen one to challenge the threat. The hero embarks on an expansive quest across diverse kingdoms and magical landscapes, recruiting fellow adventurers to aid their cause while working to restore the stolen visages and liberate the land's population from tyrannical enslavement.