Following the events of the previous autumn, Yuzuki has settled into life in the Western-style town of Yunagicho alongside the animal-eared girls of Cafe Setaria. When he helps a lost rabbit-eared girl find her sister's traditional sweets shop, he soon finds himself working at both the cake shop Sweet Tail and the newly opened Mochizuki, navigating a life that keeps getting fuller and more lively by the day.